He is the son of Will Anthonio Zeppeli and father of Caesar Anthonio Zeppeli, who plans to stop the Pillar Men. Unlike Caesar, Mario had known since he was young of the fate of his father, Will Anthonio Zeppeli. Because of this, he had researched the Stone Masks, and was able to find the Pillar Men long before Germany's discovery. Since then, he had been traveling the world in order to discover a way to destroy the creatures that were in the wall. He has five children with Caesar as the eldest.