After the defeat of his uncle Andross, Andrew Oikonny assumes leadership of the planet Venom's remaining troops and begins a rebellion. General Pepper orders an attack on Oikonny's forces, hiring the Star Fox team to assist the Cornerian Army. During the battle, Oikonny's flagship is destroyed by an insectoid creature, which attacks the Star Fox team. After Fox destroys the creature, he recovers its damaged Core Memory unit and turns it over to Beltino for research. Beltino explains the creature