"Ghost Princess" Perona[7] is the former commander of the Wild Zombies and Surprise Zombies of Thriller Bark prior to its collapse.[3] She was one of the Mysterious Four[2] and one of the major antagonists of the Thriller Bark Saga.

After the defeat of Gecko Moria's army and during the timeskip, she begrudgingly helped[8] and cooperated with Roronoa Zoro while in Dracule Mihawk's home on Kuraigana Island.[9] After the two-year timeskip, Perona has a more mature look than before, wearing a long black strapless dress with lavender frills, red high heeled boots and a big black top hat with white floral decor. She has a pink bat tattoo on her left bicep and a watch on her left wrist and wears pink nail polish on her fingernails. She still has her signature parasol, and also carries a stuffed animal resembling Kumashi with her. Additionally, she now wears her hair in multiple spiral braids on the back of her head.[12]

While reading about the Straw Hats' actions on Dressrosa and resulting new bounties, she wore a collared shirt with short, ruffled sleeves and a short dark tie hung loosely around her neck. She wore short frilly skirt with a line of crosses. She had short boots with bows on the front. She also had a large straw hat with a bow and large crown on top of it.[13] Perona followed her master, Gecko Moria, for at least ten years.[11] Her loyalty to him is somewhat complex: when she sensed defeat, she attempted to leave and abandon him[15] while noting that she had only stayed with him for fun,[31] but tried to take Bartholomew Kuma down as a final gift for Moria.[32] Despite not clearly showing it, she is said to love Moria like a father since he found her and raised her as a child, and when the newspaper falsely reported that Moria died in the Whitebeard War, Perona was deeply saddened.[33][34] When she learned that Moria is alive two years later, Perona is overjoyed, and she quickly departed Mihawk's castle to go find him.[35]

Perona had a mixed relationship with her teddy bear servant, Kumashi.